Swallowtails (Papilionidae)

• Papilio dardanus ^*
• Papilio oribazus
• Papilo epiphorbas
• Papilo delalandei
• Papilo demodocus ^*
• Graphium cyrnus
• Graphium endochus

Whites and Sulfurs (Pieridae)

• Catopsilia thaurum
• Eurema brigitta pulchella ?
• Eurema floricola floricola
• Eurema desjardinisi
• Appias spp.
• Mylothis phileris
• Mylothris smithii
• Leptosia spp.

Brush-footed Butterflies (Nymphalidae)

• Danaus chrysippus
• Charaxes cacuthis
• Charaxes andranodorus
• Charaxes analava
• Charaxes antambalou
• Charaxes cowani
• Charaxes zoolina betsimisaraka
• Euxanthe madagascariensis ^**
• Hypolimnas bolina
• Hypolimnas dexithea
• Salamis anteva
• Salamis dupreil
• Precis (= Junonia) oenone epiclelia
• Precis eurodoce
• Precis goudoti
• Precis andremiaja
• Vanessa cardui
• Eurytela dryope
• Sallya madagascariensis
• Cyrestis camillus elegans
• Neptis saclava saclava
• Aterica rabena
• Pseudacrea lucretia
• Pseudacrea glaucina
• Acraea hova
• Acraea igati
• Acraea fornax
• Acraea masamba
• Acraea eponina ?
• Acraea zitja
• Acraea encedon
• Gnophodes betsimena betsimena
• Melatistisleda
• Henotesia spp.
• Henotesia vola
• Henotesia subsimilis
• Henotesia angulifascia
• Henotesia strigula ?
• Henotesia undulosa ?
• Henotesia wardii ?
• Henotesia uniformis ?
• Henotesia cowani ^***
• Masoura spp.
• Houlbertia spp.
• Strabena spp.
• Strabena andriana ?
• Strabena smithii
• Strabena batesii
• Strabena rakoto ?
• Strabena tamatave ?
• Strabena triopthalma
• Admiratio paradoxa
• Heteropsis drepana

Metalmarks (Riodinidae)

• Saribia spp.

Blues (Lycaenidae)

• Trichiolaus mermeros
• Leptomyrina phidias ?
• Cacyreus darius ?
• Leptotes rabenafer ?
• Eichochrysops sanguigutta ?

Skippers (Hesperiidae)

• Coeliades fervida
• Celaenorrhinus humbloti
• Tagiades insularis
• Eagris sabadius
• Fulda coroller
• Galerga hyposticta
• Galerga ellipsis ?
• Hovala pardalina
• Hovala dispar
• Hovala saclavus
• Hovala arota
• Hovala amena
• Borbo spp.

Collected by Kremen, Van Epps and Rasabo between 20 July 1988 and 20 Nov 1988.
Except where noted collections were made at Talatakely (elevation 1030m).
^*Village of Ranomafana (600m)
^** Vatoharanana (1100m)
^*** approximately 2km north, northwest of Vohiparara (1300m)

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Photographs Copyright © 1998 & 1999 by George Williams